Jean-Baptiste Mallye
Université Bordeaux, Laboratoire Pacea, Faculty Member
- Archaeology, Prehistoric Archaeology, Zooarchaeology, Ethnoarchaeology, Palaeolithic Archaeology, Taphonomy, and 22 moreSmall carnivores, foxes, Small Game, Scat Analysis of Carnivores, Actualistic Studies in Taphonomy, Late Glacial Archaeology, Large Rodents, Cuon Alpinus, Human-Dog Relations, Experimental Archaeology, Vertebrate Palaeontology, Middle Palaeolithic, Aurignacian, Quaternary palaeontology, Bone diagenesis and taphonomy, Upper Palaeolithic boundary, Chatelperronian, Carnivore Taphonomy, Small Carnivores, Cutmarks, Carnivore Den, Anthropology, and Archaeological Method & Theoryedit
Les groupes culturels de la transition Pléistocène-Holocène entre Atlantique et Adriatique Actes de la séance de la Société préhistorique française de Bordeaux, 24-25 mai 2012
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"This paper presents the results of a zooarchaeological analysis conducted on the remains of large carnivores and small game from the Early Aurignacian from the site of La Quina aval. This site was excavated successively by Dr. Henri... more
"This paper presents the results of a zooarchaeological analysis conducted on the remains of large carnivores and small game from the Early Aurignacian from the site of La Quina aval. This site was excavated successively by Dr. Henri Martin and her daughter Germaine at the early and mid-twentieth century. Recently, the site was re-excavated by Véronique Dujardin. The different excavations yielded a large archaeological sample composed of many faunal remains but also lithic and bone artefact industry, a large number of personal ornament (bones, teeth and shell) as well as several anatomically modern human remains.
Our study focuses on small-game remains found during the last excavation campaign. Our results provide several unpublished results. We first offer the first quantification of each taxa in terms of number of identified remains but also in terms of minimal number of individuals.
"
Our study focuses on small-game remains found during the last excavation campaign. Our results provide several unpublished results. We first offer the first quantification of each taxa in terms of number of identified remains but also in terms of minimal number of individuals.
"
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The site of Unikoté, located in Iholdy in the French Pyrénées-Atlantiques, was discovered in 1984 and exca- vated between 1993 and 2003. The excavations were first carried out inside the cave (Unikoté I), then extended from 1995 on in... more
The site of Unikoté, located in Iholdy in the French Pyrénées-Atlantiques, was discovered in 1984 and exca- vated between 1993 and 2003. The excavations were first carried out inside the cave (Unikoté I), then extended from 1995 on in front of the entrance (Unikoté II). Both loci soon yielded traces of human occupation as well as human remains. Unikoté I has a 3-metre thick very complex stratigraphy. The artefacts found have allowed the occupation to be dated to the Middle Palaeolithic, without any greater precision. The fill of Unikoté I can be divided into three units: at the base is a hyena den with a few traces of human occupation; the intermediate level has yielded most of the human remains and the upper level is a mix of both underlying levels. The artefacts from the lower level can be attributed to the Mousterian period; those coming from the intermediate level could date from either the Upper Palaeolithic or the Mesolithic, any precise attribution to one period or the other being impossible.
Both loci have also yielded a small assemblage of human remains. In both cases the remains are extremely disturbed, which makes it impossible to reconstitute the original mode of deposit. They also lack any cultural context, so that their dating can only come from radiocarbon analyses. The Unikoté I remains date from the beginning of the Middle Neolithic II. The Unikoté II remains belong to an early phase of the Second Mesolithic. The human assemblage from the first locus is composed of 15 fragments corresponding to a minimum number of 2 individuals, a female adult and a child aged about 10. Eighty-five remains come from the second locus, among which eleven certainly belong to the same adult whose sex and age cannot be determined. The 74 remaining immature bones most probably belong to two individuals: while the corresponding frequency MNI is only 1, their stages of development are incompatible. A cranium could thus belong to a 15-year old juvenile whereas the other post-cranial remains probably belong to a child aged between 6 and 11.
The main interest of these human remains lies in the fact that they all display anthropogenic bone modifications. In the Unikoté I Neolithic assemblage the cranium shows several cutmarks that could attest to skinning practices, and a cut- mark is also visible on a fragment of scapula. The Mesolithic immature cranium from Unikoté II presents a perforation with characteristics unequivocally suggesting a perimortem trauma indicating a violent death. Moreover, several cut- ting and scraping marks can be observed on this fragment. The small size of the assemblages and their lack of context make the interpretation of these marks and the signification of these human deposits difficult to establish; while several hypotheses can be raised, none can be favoured. For Unikoté I, the specificities of the cutmarks suggest head-skinning and thus allow several possibilities to be raised: cannibalism, which cannot be proved but cannot be set aside either; a funerary practice in which the body could have been cut up but not eaten; or some specific treatment of the head, for instance its preparation to become a trophy or a relic. In Unikoté II the marks on the cranium seem to correspond more to defleshing activity, i.e. a cleaning of the bone, and the trauma matches an overall context of armed violence. It is however impossible to determine either the environment or the mobilization level of this violence, all the more so as all attested perimortem cranial traumata for the Mesolithic period are so far linked to the specific practice of head deposit known in Alsace and Southwestern Germany, which does not seem to be the case here, and as the practice of cutting up corpses is far from being rare for this period, although certainly diversely motivated.
The observations made on the human remains from Unikoté are difficult and limited, but they can nevertheless provide new information on the treatment of the dead and related body-cutting practices during the Mesolithic and Neolithic periods. Data are still lacking, so this topic cannot be discussed in detail now, until attested cases become more numer- ous. On the other hand, these observations prove once again that anthropogenic bone modifications are frequently observed and must be systematically sought in all human assemblages, whatever the period.
Both loci have also yielded a small assemblage of human remains. In both cases the remains are extremely disturbed, which makes it impossible to reconstitute the original mode of deposit. They also lack any cultural context, so that their dating can only come from radiocarbon analyses. The Unikoté I remains date from the beginning of the Middle Neolithic II. The Unikoté II remains belong to an early phase of the Second Mesolithic. The human assemblage from the first locus is composed of 15 fragments corresponding to a minimum number of 2 individuals, a female adult and a child aged about 10. Eighty-five remains come from the second locus, among which eleven certainly belong to the same adult whose sex and age cannot be determined. The 74 remaining immature bones most probably belong to two individuals: while the corresponding frequency MNI is only 1, their stages of development are incompatible. A cranium could thus belong to a 15-year old juvenile whereas the other post-cranial remains probably belong to a child aged between 6 and 11.
The main interest of these human remains lies in the fact that they all display anthropogenic bone modifications. In the Unikoté I Neolithic assemblage the cranium shows several cutmarks that could attest to skinning practices, and a cut- mark is also visible on a fragment of scapula. The Mesolithic immature cranium from Unikoté II presents a perforation with characteristics unequivocally suggesting a perimortem trauma indicating a violent death. Moreover, several cut- ting and scraping marks can be observed on this fragment. The small size of the assemblages and their lack of context make the interpretation of these marks and the signification of these human deposits difficult to establish; while several hypotheses can be raised, none can be favoured. For Unikoté I, the specificities of the cutmarks suggest head-skinning and thus allow several possibilities to be raised: cannibalism, which cannot be proved but cannot be set aside either; a funerary practice in which the body could have been cut up but not eaten; or some specific treatment of the head, for instance its preparation to become a trophy or a relic. In Unikoté II the marks on the cranium seem to correspond more to defleshing activity, i.e. a cleaning of the bone, and the trauma matches an overall context of armed violence. It is however impossible to determine either the environment or the mobilization level of this violence, all the more so as all attested perimortem cranial traumata for the Mesolithic period are so far linked to the specific practice of head deposit known in Alsace and Southwestern Germany, which does not seem to be the case here, and as the practice of cutting up corpses is far from being rare for this period, although certainly diversely motivated.
The observations made on the human remains from Unikoté are difficult and limited, but they can nevertheless provide new information on the treatment of the dead and related body-cutting practices during the Mesolithic and Neolithic periods. Data are still lacking, so this topic cannot be discussed in detail now, until attested cases become more numer- ous. On the other hand, these observations prove once again that anthropogenic bone modifications are frequently observed and must be systematically sought in all human assemblages, whatever the period.
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This papers deals with the analysis of Eurasian badger (Meles meles) remains in an archaeological context. Eurasian badgers dig large burrows as living structures and so the identification of their remains in archaeological context... more
This papers deals with the analysis of Eurasian badger (Meles meles) remains in an archaeological context. Eurasian badgers dig large burrows as living structures and so the identification of their remains in archaeological context appears doubtful. More than 400 remains of these small carnivores were recovered from Unikoté Cave. This site is assumed to be a hyena den with the occurrence of human remains and lithic artefacts. In this paper, we aimed to: 1) explain how and why so many remains of these carnivores are recognised at Unikoté Cave; 2) evaluate the role of Eurasian badgers in site formation processes and 3) to test the archaeological relevance of a bioturbated site.
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The identification of Badger remains in fossil sites raises the question over their contemporaneousness with other artefacts. Nowadays few data are available to characterize the accumulation of Badger remains in natural or archaeological... more
The identification of Badger remains in fossil sites raises the question over their contemporaneousness with other artefacts. Nowadays few data are available to characterize the accumulation of Badger remains in natural or archaeological sites. This paper aims to overcome this lack of data. We take into account three modern and five fossil samples in order to characterize the different accumulations of Badger remains. The study of three modern samples proves that the large Badger sample from Scladina cave is a natural accumulation. In Unikoté cave, large carnivores such as Hyena accumulate a part of badger remains. Accumulation by hunter-gatherer is demonstrated in Bois-Ragot cave, Faustin shelter, Unikoté cave and Rouffignac. The obtained results deliver an interpretative lens for archaeologists and taphonomists who aimed to test the homogeneity of bone samples.
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This paper focuses on modern faunal assemblages created by small carnivores in order to identify the biases they may introduce into the archaeological record. Actualistic taphonomic studies concerning the impact of large carnivores... more
This paper focuses on modern faunal assemblages created by small carnivores in order to identify the biases they may introduce into the archaeological record. Actualistic taphonomic studies concerning the impact of large carnivores (hyena, bear and wolf) on archaeological assemblages are much more developed than for small carnivores (fox and badger), yet these smaller animals are also likely to have competed with humans for sheltered spaces.
In order to study small carnivore bone accumulation behaviours from an archaeological perspective, we excavated three modern assemblages deposited by fox and badger. These allow us to explore the variability in assemblage composition relating to the season of occupation, food acquisition, bone modifications and spatial organization of the dens. Fox and badger exhibit three modes of food acquisition: hunting, scavenging and pilfering. The hunting activities of these species mainly involve small prey and the identified traces (tooth and digestion marks) are easily distinguishable from traces left by human consumption. However, the scavenging and pilfering activities of these animals include some species that are comparable in size to those hunted by humans.
Our examination of the diversity and variability in the food acquisition behaviours of small carnivores permits us to outline a number of ways in which these animals may alter archaeological assemblages. The relative abundances of different taxa and their body part representation, as well as the spatial organization of skeletal remains, tooth mark frequencies and occupation seasonality, are sources of data that are essential for detecting the activities of small carnivores at archaeological sites. Our actualistic data, combined with taphonomic analysis and insights from zooarchaeological assemblages, allow a better understanding of the impact of small carnivores and the limits which they impose on the interpretation of mixed assemblages.
In order to study small carnivore bone accumulation behaviours from an archaeological perspective, we excavated three modern assemblages deposited by fox and badger. These allow us to explore the variability in assemblage composition relating to the season of occupation, food acquisition, bone modifications and spatial organization of the dens. Fox and badger exhibit three modes of food acquisition: hunting, scavenging and pilfering. The hunting activities of these species mainly involve small prey and the identified traces (tooth and digestion marks) are easily distinguishable from traces left by human consumption. However, the scavenging and pilfering activities of these animals include some species that are comparable in size to those hunted by humans.
Our examination of the diversity and variability in the food acquisition behaviours of small carnivores permits us to outline a number of ways in which these animals may alter archaeological assemblages. The relative abundances of different taxa and their body part representation, as well as the spatial organization of skeletal remains, tooth mark frequencies and occupation seasonality, are sources of data that are essential for detecting the activities of small carnivores at archaeological sites. Our actualistic data, combined with taphonomic analysis and insights from zooarchaeological assemblages, allow a better understanding of the impact of small carnivores and the limits which they impose on the interpretation of mixed assemblages.
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"From the Upper Palaeolithic to the beginning of the Holocene, the presence of small carnivores in archaeological context is widely interpreted as evidence for peltry activities. This hypothesis is supported by several arguments,... more
"From the Upper Palaeolithic to the beginning of the Holocene, the presence of small carnivores in archaeological context is widely interpreted as evidence for peltry activities. This hypothesis is supported by several arguments, including the skeletal-part representation, the spatial distribution of bone remains, and the nature of cut marks on bones. At the end of the Palaeolithic, the question of whether small carnivores were utilised only for their skin or for their meat as well is still under debate. It has been postulated that the consumption of small carnivore meat has been underestimated in the past. In this paper, we propose a new experimental record of bone modification on small carnivore skeletons resulting from skinning by three taxidermists. The professionals produced cut marks on all the skeletons in the sample. The skinning marks are always localised on the same bones and on similar locations on those bones. We argue that this set of reference skinning marks constitutes a new tool that will prove useful in understanding the role played by small carnivores in the subsistence strategies of Palaeolithic hunter-gatherers.
FUR-USE; SMALL CARNIVORES, CUT MARKS, TAXIDERMISTS, EXPERIMENTAL STUDY, TAPHONOMY, SUBSISTENCE STRATEGIES, EUROPEAN PALAEOLITHIC"
FUR-USE; SMALL CARNIVORES, CUT MARKS, TAXIDERMISTS, EXPERIMENTAL STUDY, TAPHONOMY, SUBSISTENCE STRATEGIES, EUROPEAN PALAEOLITHIC"
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This paper presents a critical analysis of putative anthropic modifications reported on bird bones from Fontalès and badger remains from Trou de Châleux and Trou des Nutons. It turns out that these modifications are natural bone relieves.
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A single first upper molar of a Badger was discovered during the excavation of the Château Breccia. It was among debris but very close to a Middle Pleistocene context. Its small size, proportions and morphology don’t fit with the extant... more
A single first upper molar of a Badger was discovered during the excavation of the Château Breccia. It was among debris but very close to a Middle Pleistocene context. Its small size, proportions and morphology don’t fit with the extant Meles meles. A critical overview of the genus Meles from European Plio-Pleistocene is tried as an attempt to determinate the species. The comparison with Meles thorali, M. dimitrius, M. hollitzeri, M. meles atavus, M. thorali spelaeus don’t permit to attribute this molar to any of these species. Some of them are well defined in space and time, the others should be worth reviewed. Finally, the molar from Château is refered to Meles sp., pointing to a small sized animal obviously in the lineage of Meles meles.
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"Jaurens Cave yielded four Mustelid species, the wolverine Gulo gulo, the steppe polecat Mustela (Putorius) eversmanni, the stoat Mustela erminea and the weasel Mustela nivalis, all and especially the two first ones being rare in the... more
"Jaurens Cave yielded four Mustelid species, the wolverine
Gulo gulo, the steppe polecat Mustela
(Putorius) eversmanni, the stoat Mustela erminea and
the weasel Mustela nivalis, all and especially the two
first ones being rare in the french Upper Pleistocene.
The wolverine with 60 remains, including two complete
skulls, is the most numerous Mustelid. Weighing
about 35 kilograms, it is biometrically much larger
than the recent specimens of the same species. The
steppe polecat, the stoat and the weasel are very similar
in size to the recent ones,
The occurence of these four Mustelids shows a kind of
palaeoenvironment under a steppic dominance but
with a mosaic of small forested zones, and the temperature
was not especially cold."
Gulo gulo, the steppe polecat Mustela
(Putorius) eversmanni, the stoat Mustela erminea and
the weasel Mustela nivalis, all and especially the two
first ones being rare in the french Upper Pleistocene.
The wolverine with 60 remains, including two complete
skulls, is the most numerous Mustelid. Weighing
about 35 kilograms, it is biometrically much larger
than the recent specimens of the same species. The
steppe polecat, the stoat and the weasel are very similar
in size to the recent ones,
The occurence of these four Mustelids shows a kind of
palaeoenvironment under a steppic dominance but
with a mosaic of small forested zones, and the temperature
was not especially cold."
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Faunal remains from spoil heaps of two burrows inhabited by small carnivores (fox and badger) are analysed from a taphonomical point of view. This analysis provides characteristics for bone accumulation produced by small carnivores and... more
Faunal remains from spoil heaps of two burrows inhabited by small carnivores (fox and badger) are analysed from a taphonomical point of view. This analysis provides characteristics for bone accumulation produced by small carnivores and will be a powerful tool for deciphering site formation about the occupational alternation of small carnivores and humans. Identified species were grouped by size classes. Faunal spectrum is composed by varied species of microfauna (70%), mesofauna (30%). Macromammal remains are under represented (less than 1%) and come from scavenged carcasses. Predators and consumed species are compared on the basis of the skeletal part representation, age classes and recording of predation marks (gnawed and digested bones). Skeletal part representation shows that all taxons exhibit a low-representation of axial skeleton and autopodial bones. Predators show a high representation of hind limb bones and a low representation of the fore limb bones whereas consumed species exhibit a reverse pattern. Mortality curve analysis provides an attritional profile for carnivores and helps for the establishment of the season of occupation of the burrow. Adults largely dominate consumed species. Moreover, predation marks are found in 1% of the carnivores’ bones and from 15% to more than 40% on prey bones. The large-sized prey bones only wear gnawed marks (20%) and anthropic marks (10–20%) whereas microfaunal remains exhibits more digested marks (40%) than gnawing stigmata (5%). Medium-sized animals bones wear both marks and with the same proportions (10–15%).
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"Fur exploitation is usually advocate when Small carnivores remains are identified in archaeological sites. However, few butchery referential are available to test this assumption. An experimental study was conduced in order to identify... more
"Fur exploitation is usually advocate when Small carnivores remains are identified in archaeological sites. However, few butchery referential are available to test this assumption. An experimental study was conduced in order to identify cut marks derived from skinning activities on small carnivores carcasses. Several skinning methods were tested. Results show that cut mark location varies according to the skinning method employed. These methods provide a new insight on skeletal part representation regarding site function (home basement, kill site, pelts site) identified in archaeological record. Moreover, this study provides new tools to discuss about the status of these small games in human societies.
Keywords: Experimental study, cut marks, fur, Small carnivores, skinning"
Keywords: Experimental study, cut marks, fur, Small carnivores, skinning"
Research Interests:
Research Interests:
Understanding the integrity of archaeological accumulations and any taphonomic process that interfered with the integrity of accumulations is necessary to assess hominid behaviour and ecology. From this point of view, the occurrence... more
Understanding the integrity of archaeological accumulations and any taphonomic process that interfered with the integrity of accumulations is necessary to assess hominid behaviour and ecology. From this point of view, the occurrence within archaeological sites of large sized burrowers, such as badgers, could be problematic. Badger burrows and skeletal remains frequently occur in Pleistocene deposits; however, the temporal and behavioural relationships of these items to the rest of the archaeological assemblage are frequently unknown, due to a poor understanding of badger species—either from the palaeontological view or from its relationships with hominids. Thus, a part of our work is devoted to a synthesis concerning 1) badger phylogeny, 2) criteria for diagnosing the accumulation of burrowing animal remains, 3) the exploitation of carnivores by humans during the Palaeolithic. Moreover, a part of this work is dedicated to understanding badger induced bioturbation on archaeological sites. Neotaphonomical experiments were run in natural contexts and butchery experiments were conducted on small carnivores carcasses. These methodological tools were then applied to archaeological material from Scladina Cave in Belgium (Middle Palaeolithic). In France, material analysed comes from Artenac (Charente, Mousterian), Bois-Ragot (Vienne, Azilian), Faustin shelter (Gironde, Late Magdalenian), Rouffignac (Dordogne, Mesolithic) and Unikoté (Pyrenées-Atlantique, Middle Palaeolithic till actual). In all these sites the origins of the badger remains were clearly identified and then we discuss the validity of the archaeological assemblages. In the sites for which an anthropic origin was demonstrated, we conducted zooarchaeological analyses. Our results suggest that humans have been exploiting carnivores since the Middle Palaeolithic.
mots-clés en anglais : Eurasian badger – Meles – Taphonomy – Neotaphonomy – Zooarchaeology – Experimentation – Palaeolithic – Plio-Pleistocene – Palaeontology – Bioturbation
mots-clés en anglais : Eurasian badger – Meles – Taphonomy – Neotaphonomy – Zooarchaeology – Experimentation – Palaeolithic – Plio-Pleistocene – Palaeontology – Bioturbation
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"Biostratigraphic and paleoecological analyses of Southeast Asian Pleistocene faunal sites are based on the assumption that paleontological assemblages are homogeneous. This means that the sites formed during a time range shorter than... more
"Biostratigraphic and paleoecological analyses of Southeast Asian Pleistocene faunal sites are based on the assumption that paleontological assemblages are homogeneous. This means that the sites formed during a time range shorter than biological evolution range and, above all, without faunal replacement induced by environmental successions. Detailed study of the Cave of the Monk paleontological site, in Thailand, has lead to the conclusion that the complex pattern of this Southeast Asian paleontological site cannot support such an assumption.
The Cave of the Monk provides an Ailuropoda–Stegodon assemblage typical of Southeast Asia Pleistocene Fauna. The pluridisciplinary study presented here includes (i) site morphology description, (ii) sedimentological analysis of fossiliferous deposits, (iii) taphonomy study of the bone assemblage, and (iv) electron spin resonance (ESR) dating of tooth samples. This integrated approach demonstrates the existence of a Pleistocene porcupine den inhabited during MIS 3 and 2. The features of this site are commonly reported for other paleontological sites of the area, indicating that this kind of site formation is a very general one in Southeast Asia.
ESR dating and porcupine ethological reports are used to question the site time averaging. A time range of a few tens of thousands of years is suggested. The paleoecological value of the site can then be discussed. On one hand, the assemblage is thought to be a valuable paleoenvironmental record considering that the bone accumulation agent is reputed to represent the fauna present in the environment without bias. On the other hand, the time averaging of the site indicates that site formation range likely covered the environmental fluctuations of the last climatic cycle. This last hypothesis is confirmed by the alternating association of species indicated by the stratigraphic distribution of the faunal remains.
The research demonstrates the need for integrated and pluridisciplinary taphonomic study of Southeast Asian paleontological sites. It also reveals the utility of a microstratigraphic analysis of a fossiliferous karst deposit in order to disentangle the succession of Pleistocene mammals in response to environmental changes."
The Cave of the Monk provides an Ailuropoda–Stegodon assemblage typical of Southeast Asia Pleistocene Fauna. The pluridisciplinary study presented here includes (i) site morphology description, (ii) sedimentological analysis of fossiliferous deposits, (iii) taphonomy study of the bone assemblage, and (iv) electron spin resonance (ESR) dating of tooth samples. This integrated approach demonstrates the existence of a Pleistocene porcupine den inhabited during MIS 3 and 2. The features of this site are commonly reported for other paleontological sites of the area, indicating that this kind of site formation is a very general one in Southeast Asia.
ESR dating and porcupine ethological reports are used to question the site time averaging. A time range of a few tens of thousands of years is suggested. The paleoecological value of the site can then be discussed. On one hand, the assemblage is thought to be a valuable paleoenvironmental record considering that the bone accumulation agent is reputed to represent the fauna present in the environment without bias. On the other hand, the time averaging of the site indicates that site formation range likely covered the environmental fluctuations of the last climatic cycle. This last hypothesis is confirmed by the alternating association of species indicated by the stratigraphic distribution of the faunal remains.
The research demonstrates the need for integrated and pluridisciplinary taphonomic study of Southeast Asian paleontological sites. It also reveals the utility of a microstratigraphic analysis of a fossiliferous karst deposit in order to disentangle the succession of Pleistocene mammals in response to environmental changes."
Research Interests:
... 5 : Laboratoire des sciences du climat et de l'environnement (LSCE). CNRS : UMR1572 INSU CEA : DSM/LSCE Université de Versailles-Saint Quentin en Yvelines Institut... more
... 5 : Laboratoire des sciences du climat et de l'environnement (LSCE). CNRS : UMR1572 INSU CEA : DSM/LSCE Université de Versailles-Saint Quentin en Yvelines Institut Pierre Simon Laplace. 6 : Geophysical institute. Bulgarian Academy of Sciences. ...
Research Interests:
Research Interests:
In 2002, a Neandertal partial femoral diaphysis was discovered at Les Rochers-de-Villeneuve (Vienne, France). Radiocarbon dated to +/- 40,700 14C years before present, this specimen is one of the most recent Middle Paleolithic... more
In 2002, a Neandertal partial femoral diaphysis was discovered at Les Rochers-de-Villeneuve (Vienne, France). Radiocarbon dated to +/- 40,700 14C years before present, this specimen is one of the most recent Middle Paleolithic Neandertals. The diaphysis derives from an archeological level indicating alternating human and carnivore (mostly hyena) occupation of the cave, reinforcing the close proximity and probable competition of Middle Paleolithic humans with large carnivores for resources and space. Morphological aspects of the diaphysis and ancient DNA extracted from it indicate that it is aligned with the Neandertals and is distinct from early modern humans. However, its midshaft cortical bone distribution places it between other Middle Paleolithic Neandertals and the Châtelperronian Neandertal from La Roche-à-Pierrot, supporting a pattern of changing mobility patterns among late Middle Paleolithic Neandertals on the eve of modern human dispersals into Europe.
